About the Authors

Paul G. Matson

pmatson@gmail.com

Affiliation Department of Ecology, Evolution, and Marine Biology, University of California Santa Barbara, Santa Barbara, California, United States of America

Libe Washburn

Affiliation Department of Geography, University of California Santa Barbara, Santa Barbara, California, United States of America

Todd R. Martz

Affiliation Geosciences Research Division, Scripps Institution of Oceanography, University of California San Diego, La Jolla, California, United States of America

Gretchen E. Hofmann

Affiliation Department of Ecology, Evolution, and Marine Biology, University of California Santa Barbara, Santa Barbara, California, United States of America

Competing Interests

Dr. Gretchen Hofmann is an editorial board member at PLOS ONE but that does not alter the authors’ adherence to PLOS ONE editorial policies and criteria.

Author Contributions

Conceived and designed the experiments: PGM GEH. Performed the experiments: PGM GEH. Analyzed the data: PGM GEH LW TRM. Contributed reagents/materials/analysis tools: GEH LW. Contributed to the writing of the manuscript: PGM GEH LW TRM.