About the Authors

Yutaka Terada

Affiliation Laboratory of Veterinary Microbiology, Joint Faculty of Veterinary Medicine, Yamaguchi University, Yamaguchi, Japan

Nobutaka Matsui

Affiliation Laboratory of Veterinary Microbiology, Joint Faculty of Veterinary Medicine, Yamaguchi University, Yamaguchi, Japan

Keita Noguchi

Affiliation Laboratory of Veterinary Microbiology, Joint Faculty of Veterinary Medicine, Yamaguchi University, Yamaguchi, Japan

Ryusei Kuwata

Affiliation Laboratory of Veterinary Microbiology, Joint Faculty of Veterinary Medicine, Yamaguchi University, Yamaguchi, Japan

Hiroshi Shimoda

Affiliation Laboratory of Veterinary Microbiology, Joint Faculty of Veterinary Medicine, Yamaguchi University, Yamaguchi, Japan

Takehisa Soma

Affiliation Veterinary Diagnostic Laboratory, Marupi Lifetech Co. Ltd., Osaka, Japan

Masami Mochizuki

Affiliation Laboratory of Emerging Infectious Diseases, Joint Faculty of Veterinary Medicine, Kagoshima University, Kagoshima, Japan

Ken Maeda

kmaeda@yamaguchi-u.ac.jp

Affiliation Laboratory of Veterinary Microbiology, Joint Faculty of Veterinary Medicine, Yamaguchi University, Yamaguchi, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KM. Performed the experiments: YT NM KN RK HS TS MM KM. Analyzed the data: YT KM. Contributed reagents/materials/analysis tools: MM TS KM. Contributed to the writing of the manuscript: YT NM KM.