About the Authors

Yoko Hayashi-Takanaka

Affiliations Graduate School of Frontier Biosciences, Osaka University, Suita, Japan, Graduate School of Bioscience and Biotechnology, Tokyo Institute of Technology, Yokohama, Japan

Timothy J. Stasevich

Affiliations Graduate School of Frontier Biosciences, Osaka University, Suita, Japan, Department of Biochemistry and Molecular Biology, Colorado State University, Fort Collins, Colorado, United States of America, Janelia Farm Research Campus, Howard Hughes Medical Institute, Ashburn, Virginia, United States of America

Hitoshi Kurumizaka

Affiliation Graduate School of Advanced Science and Engineering, Waseda University, Tokyo, Japan

Naohito Nozaki

Affiliation Mab Institute Inc, Sapporo, Japan

Hiroshi Kimura

hkimura@bio.titech.ac.jp

Affiliations Graduate School of Frontier Biosciences, Osaka University, Suita, Japan, Graduate School of Bioscience and Biotechnology, Tokyo Institute of Technology, Yokohama, Japan

Competing Interests

N. Nozaki is a founder of MAB Institute Inc. and the salary is paid by the company. This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: YH-T H. Kimura. Performed the experiments: YH-T. Analyzed the data: YH-T. Contributed reagents/materials/analysis tools: TJS H. Kurumizaka NN. Contributed to the writing of the manuscript: YH-T H. Kimura.