About the Authors

Jiankai Wei

Affiliations Key Laboratory of Experimental Marine Biology, Institute of Oceanology, Chinese Academy of Sciences, Qingdao, China, University of Chinese Academy of Sciences, Beijing, China

Xiaojun Zhang

Affiliation Key Laboratory of Experimental Marine Biology, Institute of Oceanology, Chinese Academy of Sciences, Qingdao, China

Yang Yu

Affiliations Key Laboratory of Experimental Marine Biology, Institute of Oceanology, Chinese Academy of Sciences, Qingdao, China, University of Chinese Academy of Sciences, Beijing, China

Hao Huang

Affiliation Hainan Guandtop Ocean Breeding Co. Ltd, Haikou, China

Fuhua Li

Affiliation Key Laboratory of Experimental Marine Biology, Institute of Oceanology, Chinese Academy of Sciences, Qingdao, China

Jianhai Xiang

jhxiang@qdio.ac.cn

Affiliation Key Laboratory of Experimental Marine Biology, Institute of Oceanology, Chinese Academy of Sciences, Qingdao, China

Competing Interests

The authors have the following interests: Co-author Hao Huang is employed by Hainan Guandtop Ocean Breeding Co. Ltd. Hainan Guandtop Ocean Breeding Co. Ltd provided shrimp samples used in this study. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: XZ FL JX. Performed the experiments: JW XZ YY. Analyzed the data: JW XZ. Contributed reagents/materials/analysis tools: YY HH. Contributed to the writing of the manuscript: JW XZ FL JX.