About the Authors

Hiroshi Katoh

katohh@affrc.go.jp

Affiliation NARO Institute of Fruit Tree Science, Tsukuba, Ibaraki, Japan

Shin-ichi Miyata

Affiliation NARO Institute of Fruit Tree Science, Tsukuba, Ibaraki, Japan

Hiromitsu Inoue

Affiliation Kuchinotsu Citrus Research Station, NARO Institute of Fruit Tree Science, Minami-shimabara, Nagasaki, Japan

Toru Iwanami

Affiliation NARO Institute of Fruit Tree Science, Tsukuba, Ibaraki, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TI. Performed the experiments: HK SM HI. Analyzed the data: HK SM. Contributed reagents/materials/analysis tools: SM HI TI. Contributed to the writing of the manuscript: HK SM HI TI.