About the Authors

Mathieu Andraud

mathieu.andraud@anses.fr

Affiliations Pig Epidemiology and Welfare Unit, Anses, Laboratoire de Ploufragan-Plouzané, Ploufragan, France, Université Européenne de Bretagne, Rennes, France

Maribel Casas

Affiliations Centre for Research in Environmental Epidemiology (CREAL), Barcelona, Spain, Centre de Recerca en Sanitat Animal (CReSA), Barcelona, Spain

Nicole Pavio

Affiliations UMR 1161 Virology, Anses, Laboratoire de Santé Animale, Maisons-Alfort, France, UMR 1161 Virology, INRA, Maisons-Alfort, France, UMR 1161 Virology, Université Paris-Est, Ecole Nationale Vétérinaire d′Alfort, Maisons-Alfort, France

Nicolas Rose

Affiliations Pig Epidemiology and Welfare Unit, Anses, Laboratoire de Ploufragan-Plouzané, Ploufragan, France, Université Européenne de Bretagne, Rennes, France

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: NR MA NP. Performed the experiments: MA NR NP. Analyzed the data: MA MC. Contributed reagents/materials/analysis tools: MC NP. Wrote the paper: MA NR NP MC.