About the Authors

Yosuke Matsumoto

Affiliation Department of Molecular Physiology, College of Pharmaceutical Sciences, Ritsumeikan University, Kusatsu, Shiga, Japan

Masatoshi Inden

Affiliation Laboratory of Medical Therapeutics and Molecular Therapeutics, Gifu Pharmaceutical University, Gifu, Japan

Atsushi Tamura

Affiliation Laboratory of Biological Science, Graduate School of Frontier Biosciences and Graduate School of Medicine, Osaka University, Suita, Osaka, Japan

Ryo Hatano

Affiliation Department of Molecular Physiology, College of Pharmaceutical Sciences, Ritsumeikan University, Kusatsu, Shiga, Japan

Sachiko Tsukita

Affiliation Laboratory of Biological Science, Graduate School of Frontier Biosciences and Graduate School of Medicine, Osaka University, Suita, Osaka, Japan

Shinji Asano

ashinji@ph.ritsumei.ac.jp

Affiliation Department of Molecular Physiology, College of Pharmaceutical Sciences, Ritsumeikan University, Kusatsu, Shiga, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YM SA. Performed the experiments: YM AT. Analyzed the data: YM RH SA MI. Contributed reagents/materials/analysis tools: ST AT. Contributed to the writing of the manuscript: YM SA.