About the Authors

Erin Manis

Affiliation Department of Biological Sciences, Kent State University, Kent, Ohio, United States of America

Todd V. Royer

Affiliation School of Public and Environmental Affairs, Indiana University, Bloomington, Indiana, United States of America

Laura T. Johnson

Current address: National Center for Water Quality Research, Heidelberg University, Tiffin, Ohio, United States of America

Affiliation School of Public and Environmental Affairs, Indiana University, Bloomington, Indiana, United States of America

Laura G. Leff

lleff@kent.edu

Affiliation Department of Biological Sciences, Kent State University, Kent, Ohio, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: EM TR LJ LL. Performed the experiments: EM LJ. Analyzed the data: EM TR LJ LL. Contributed reagents/materials/analysis tools: TR LL. Wrote the paper: EM TR LJ LL.