About the Authors

Takashi Narihiro

t.narihiro@aist.go.jp

Affiliation Bioproduction Research Institute, National Institute of Advanced Industrial Science and Technology (AIST), Tsukuba, Ibaraki, Japan

Hideyuki Tamaki

Affiliation Bioproduction Research Institute, National Institute of Advanced Industrial Science and Technology (AIST), Tsukuba, Ibaraki, Japan

Aya Akiba

Affiliation Bioproduction Research Institute, National Institute of Advanced Industrial Science and Technology (AIST), Tsukuba, Ibaraki, Japan

Kazuto Takasaki

Affiliation FASMAC Co., Ltd., Atsugi, Kanagawa, Japan

Koichiro Nakano

Affiliation FASMAC Co., Ltd., Atsugi, Kanagawa, Japan

Yoichi Kamagata

Affiliations Bioproduction Research Institute, National Institute of Advanced Industrial Science and Technology (AIST), Tsukuba, Ibaraki, Japan, Bioproduction Research Institute, National Institute of Advanced Industrial Science and Technology (AIST), Sapporo, Hokkaido, Japan

Satoshi Hanada

Affiliation Bioproduction Research Institute, National Institute of Advanced Industrial Science and Technology (AIST), Tsukuba, Ibaraki, Japan

Taizo Maji

Affiliation Shubun University, Ichinomiya, Aichi, Japan

Competing Interests

TN, HT, AA, YK, SH and TM have neither financial nor occupational affiliations with FASMAC Co., Ltd. at the time of the study. KT and KN were employees of FASMAC Co., Ltd. at the time of the study. There are no patents, products in development or marketed products to declare. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials, as detailed online in the guide for authors.

Author Contributions

Conceived and designed the experiments: TN HT YK SH TM. Performed the experiments: TN AA. Analyzed the data: TN KT KN. Contributed reagents/materials/analysis tools: TN TM. Contributed to the writing of the manuscript: TN HT YK SH.