About the Authors

Misugi Uraji

Affiliation Research Institute for Biological Sciences (RIBS), Okayama Prefectural Technology Center for Agriculture, Forestry, and Fisheries, Kaga-gun, Okayama, Japan

Jiro Arima

Affiliation Department of Agricultural, Biological, and Environmental Sciences, Faculty of Agriculture, Tottori University, Tottori, Japan

Yoshikazu Inoue

Affiliation Satake Corporation, Higashi-Hiroshima-shi, Hiroshima, Japan

Koichi Harazono

Affiliation Enzyme Division, Bio Chemicals Department, Nagase ChemteX Corporation, Fukuchiyama, Kyoto, Japan

Tadashi Hatanaka

hatanaka@bio-ribs.com

Affiliation Research Institute for Biological Sciences (RIBS), Okayama Prefectural Technology Center for Agriculture, Forestry, and Fisheries, Kaga-gun, Okayama, Japan

Competing Interests

In addition, all authors in the manuscript do not have problems of any other relevant declarations relating to employment, consultancy, patents, products in development or marketed products, etc. This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: MU TH. Performed the experiments: MU JA. Analyzed the data: MU. Contributed reagents/materials/analysis tools: YI KH. Contributed to the writing of the manuscript: MU TH.