About the Authors

Sadegh Mousavi

Contributed equally to this work with: Sadegh Mousavi, Arghavan Alisoltani

Affiliation Department of Plant Breeding and Biotechnology, Faculty of Agriculture, Shahrekord University, Shahrekord, Iran

Arghavan Alisoltani

Contributed equally to this work with: Sadegh Mousavi, Arghavan Alisoltani

Affiliation Department of Plant Breeding and Biotechnology, Faculty of Agriculture, Shahrekord University, Shahrekord, Iran

Behrouz Shiran

shiran@agr.sku.ac.ir

Affiliations Department of Plant Breeding and Biotechnology, Faculty of Agriculture, Shahrekord University, Shahrekord, Iran, Institute of Biotechnology, Shahrekord University, Shahrekord, Iran

Hossein Fallahi

Affiliation Department of Biology, School of Sciences, Razi University, Bagh-e-Abrisham Kermanshah, Iran

Esameil Ebrahimie

Affiliations Department of Crop Production and Plant Breeding, Faculty of Agriculture, Shiraz University, Shiraz, Iran, School of Molecular and Biomedical Science, The University of Adelaide, Adelaide, Australia

Ali Imani

Affiliation Department of Horticulture, Seed and Plant Improvement Institute (SPII), Karaj, Iran

Saadollah Houshmand

Affiliation Department of Plant Breeding and Biotechnology, Faculty of Agriculture, Shahrekord University, Shahrekord, Iran

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BS SM AA EE. Performed the experiments: AA HF SM BS. Analyzed the data: HF SM AA BS EE. Contributed reagents/materials/analysis tools: BS AI SH. Wrote the paper: AA HF SM BS EE. Designed the codes used in analysis: HF SM AA BS EE.