About the Authors

Cuizhu Zhao

Affiliations The Key Laboratory of Plant Cell Engineering and Germplasm Innovation, School of Life Sciences, Shandong University, Jinan, Shandong, China, College of Agronomy, Northwest A&F University, Yangling, Shanxi, China

Jin Xu

Affiliation Key Laboratory of Agricultural Water Resources, Center for Agricultural Resources Research, Institute of Genetics and Developmental Biology, Chinese Academy of Sciences, Shijiazhuang, China

Qiang Li

Affiliation The Key Laboratory of Plant Cell Engineering and Germplasm Innovation, School of Life Sciences, Shandong University, Jinan, Shandong, China

Shuo Li

Affiliation The Key Laboratory of Plant Cell Engineering and Germplasm Innovation, School of Life Sciences, Shandong University, Jinan, Shandong, China

Peng Wang

Affiliation The Key Laboratory of Plant Cell Engineering and Germplasm Innovation, School of Life Sciences, Shandong University, Jinan, Shandong, China

Fengning Xiang

xfn0990@sdu.edu.cn

Affiliation The Key Laboratory of Plant Cell Engineering and Germplasm Innovation, School of Life Sciences, Shandong University, Jinan, Shandong, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: FX. Performed the experiments: CZ PW QL. Analyzed the data: FX CZ PW. Contributed reagents/materials/analysis tools: SL. Wrote the paper: CZ JX SL.