About the Authors

D. Richard Cameron

dcameron@tnc.org

Affiliation The Nature Conservancy, San Francisco, California, United States of America

Jaymee Marty

Current address: Marty Ecological Consulting, Sacramento, California, United States of America

Affiliation The Nature Conservancy, San Francisco, California, United States of America

Robert F. Holland

Affiliation Geobotanical Phenomenology, Auburn, California, United States of America

Competing Interests

JM is currently an employee of Marty Ecological Consulting. RH is an employee of Geobotanical Phenomenology. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: DC JM. Performed the experiments: DC RH. Analyzed the data: DC RH. Wrote the paper: DC JM RH.