About the Authors

Erica C. Nakajima

Affiliations Physician Scientist Training Program, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America, Howard Hughes Medical Fellow, Howard Hughes Medical Institute, Bethesda, Maryland, United States of America, Department of Pharmacology & Chemical Biology, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Charles Laymon

Affiliation Department of Radiology, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

Matthew Oborski

Affiliation Department of Radiology, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

Weizhou Hou

Affiliation Department of Immunology, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

Lin Wang

Affiliation Department of Pathology, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

Jennifer R. Grandis

Affiliation Department of Otolaryngology, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

Robert L. Ferris

Affiliation Department of Otolaryngology, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

James M. Mountz

Affiliation Department of Radiology, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

Bennett Van Houten

vanhoutenb@upmc.edu

Affiliation Department of Pharmacology & Chemical Biology, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Competing Interests

JRG receives funding from Novartis and BMS. None of the authors have any additional conflicts of interests and have nothing to declare with regard to the experiments presented in this study.

Author Contributions

Conceived and designed the experiments: ECN CML BVH JRG. Performed the experiments: ECN WH CML MJO. Analyzed the data: ECN CML MJO JMM BVH JRG LW. Contributed reagents/materials/analysis tools: WH JMM RLF JRG. Contributed to the writing of the manuscript: ECN CLM JMM BVH.