About the Authors

Tzung-Yi Lin

Affiliation Institute of Bioscience and Biotechnology, National Taiwan Ocean University, Keelung, Taiwan

Chi-Fu Chou

Affiliation Institute of Bioscience and Biotechnology, National Taiwan Ocean University, Keelung, Taiwan

Hsin-Yu Chung

Affiliation Institute of Bioscience and Biotechnology, National Taiwan Ocean University, Keelung, Taiwan

Chia-Yin Chiang

Affiliation Institute of Bioscience and Biotechnology, National Taiwan Ocean University, Keelung, Taiwan

Chung-Hao Li

Affiliation Institute of Bioscience and Biotechnology, National Taiwan Ocean University, Keelung, Taiwan

Jen-Leih Wu

Affiliation Institute of Cellular and Organismic Biology, Academia Sinica, Taipei, Taiwan

Han-Jia Lin

Affiliation Institute of Bioscience and Biotechnology, National Taiwan Ocean University, Keelung, Taiwan

Tun-Wen Pai

Affiliation Department of Computer Science and Engineering, National Taiwan Ocean University, Keelung, Taiwan

Chin-Hwa Hu

Affiliation Institute of Bioscience and Biotechnology, National Taiwan Ocean University, Keelung, Taiwan

Wen-Shyong Tzou

wstzou@ntou.edu.tw

Affiliation Department of Life Sciences, National Taiwan Ocean University, Keelung, Taiwan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TYL CHH WST. Performed the experiments: TYL CFC CYC. Analyzed the data: TYL HYC CHL. Contributed reagents/materials/analysis tools: JLW HJL TWP. Wrote the paper: TYL CHH WST.