About the Authors

Tomohiro Osugi

Current address: Bioorganic Research Institute, Suntory Foundation for Life Sciences, Shimamoto, Mishima, Osaka, Japan

Affiliation Laboratory of Integrative Brain Sciences, Department of Biology, Waseda University, and Center for Medical Life Science of Waseda University, Tokyo, Japan

Tomoki Okamura

Affiliation Laboratory of Integrative Brain Sciences, Department of Biology, Waseda University, and Center for Medical Life Science of Waseda University, Tokyo, Japan

You Lee Son

Affiliation Laboratory of Integrative Brain Sciences, Department of Biology, Waseda University, and Center for Medical Life Science of Waseda University, Tokyo, Japan

Makoto Ohkubo

Affiliation Laboratory of Integrative Brain Sciences, Department of Biology, Waseda University, and Center for Medical Life Science of Waseda University, Tokyo, Japan

Takayoshi Ubuka

Affiliation Laboratory of Integrative Brain Sciences, Department of Biology, Waseda University, and Center for Medical Life Science of Waseda University, Tokyo, Japan

Yasuhisa Henmi

Affiliation Aitsu Marine Station, Center for Marine Environmental Studies, Kumamoto University, Kumamoto, Japan

Kazuyoshi Tsutsui

k-tsutsui@waseda.jp

Affiliation Laboratory of Integrative Brain Sciences, Department of Biology, Waseda University, and Center for Medical Life Science of Waseda University, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: T. Osugi YLS TU YH KT. Performed the experiments: T. Osugi T. Okamura YLS MO. Analyzed the data: T. Osugi T. Okamura YLS MO TU KT. Contributed reagents/materials/analysis tools: T. Osugi YLS MO YH KT. Wrote the paper: T. Osugi YLS TU KT.