About the Authors

Manuel A. Giannoni-Guzmán

Affiliation Department of Biology, University of Puerto Rico Rio Piedras, San Juan, Puerto Rico

Tugrul Giray

Affiliation Department of Biology, University of Puerto Rico Rio Piedras, San Juan, Puerto Rico

Jose Luis Agosto-Rivera

Affiliation Department of Biology, University of Puerto Rico Rio Piedras, San Juan, Puerto Rico

Blake K. Stevison

Affiliation Laboratory of Behavioral Biology and Comparative Psychology, Oklahoma State University, Stillwater, Oklahoma, United States of America

Brett Freeman

Affiliation Laboratory of Behavioral Biology and Comparative Psychology, Oklahoma State University, Stillwater, Oklahoma, United States of America

Paige Ricci

Affiliation Laboratory of Behavioral Biology and Comparative Psychology, Oklahoma State University, Stillwater, Oklahoma, United States of America

Erika A. Brown

Affiliation Laboratory of Behavioral Biology and Comparative Psychology, Oklahoma State University, Stillwater, Oklahoma, United States of America

Charles I. Abramson

charles.abramson@okstate.edu

Affiliation Laboratory of Behavioral Biology and Comparative Psychology, Oklahoma State University, Stillwater, Oklahoma, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TG CIA. Performed the experiments: MAGG PR BS BF. Analyzed the data: MAGG EB TG. Contributed reagents/materials/analysis tools: JLAR TG CIA. Wrote the paper: JLAR TG CIA.