About the Authors

Jiangnan Luo

Affiliation Department of Zoology, Stockholm University, Stockholm, Sweden

Oleh V. Lushchak

Affiliation Department of Zoology, Stockholm University, Stockholm, Sweden

Philip Goergen

Affiliation Department of Neuroscience, Uppsala University, Uppsala, Sweden

Michael J. Williams

Affiliation Department of Neuroscience, Uppsala University, Uppsala, Sweden

Dick R. Nässel

dnassel@zoologi.su.se

Affiliation Department of Zoology, Stockholm University, Stockholm, Sweden

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JL OVL PG MJW DRN. Performed the experiments: JL OVL PG. Analyzed the data: JL OVL PG MJW DRN. Wrote the paper: JL DRN. Edited manuscript (wrote behavior parts): MJW.