About the Authors

Felicity V. Crotty

Affiliation Institute of Biological, Environmental and Rural Sciences, Aberystwyth University, Gogerddan, Aberystwyth, United Kingdom

Rhun Fychan

Affiliation Institute of Biological, Environmental and Rural Sciences, Aberystwyth University, Gogerddan, Aberystwyth, United Kingdom

Vince J. Theobald

Affiliation Institute of Biological, Environmental and Rural Sciences, Aberystwyth University, Gogerddan, Aberystwyth, United Kingdom

Ruth Sanderson

Affiliation Institute of Biological, Environmental and Rural Sciences, Aberystwyth University, Gogerddan, Aberystwyth, United Kingdom

David R. Chadwick

Affiliation Environment Centre Wales, School of Environment, Natural Resources and Geography, Bangor University, Bangor, Gwynedd, United Kingdom

Christina L. Marley

cvm@aber.ac.uk

Affiliation Institute of Biological, Environmental and Rural Sciences, Aberystwyth University, Gogerddan, Aberystwyth, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CLM RF DRC. Performed the experiments: RF VJT. Analyzed the data: FVC RS. Wrote the paper: FVC CLM DRC.