About the Authors

Heather A. Cole

Contributed equally to this work with: Heather A. Cole, Tetsuro Ohba

Affiliation Department of Orthopaedics and Rehabilitation, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America

Tetsuro Ohba

Contributed equally to this work with: Heather A. Cole, Tetsuro Ohba

Affiliation Department of Orthopaedics and Rehabilitation, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America

Jiro Ichikawa

Affiliation Department of Orthopaedics and Rehabilitation, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America

Jeffry S. Nyman

Affiliation Department of Orthopaedics and Rehabilitation, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America

Justin M. M. Cates

Affiliation Department of Pathology, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America

Hirotaka Haro

Affiliation Department of Orthopaedic Surgery, Faculty of Medicine, University of Yamanashi, Yamanashi, Japan

Herbert S. Schwartz

Affiliation Department of Orthopaedics and Rehabilitation, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America

Jonathan G. Schoenecker

jon.schoenecker@vanderbilt.edu

Affiliation Department of Orthopaedics and Rehabilitation, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HC TO JI JS. Performed the experiments: HC TO JI. Analyzed the data: HC TO JI. Contributed reagents/materials/analysis tools: HC TO JN HH HS JS. Wrote the paper: HC TO JN JC HH HS JS.