About the Authors

Kihyuck Kwak

Affiliation Department of Pathology, The Johns Hopkins University, Baltimore, Maryland, United States of America

Rosie Jiang

Affiliation Department of Pathology, The Johns Hopkins University, Baltimore, Maryland, United States of America

Joshua W. Wang

Affiliation Department of Pathology, The Johns Hopkins University, Baltimore, Maryland, United States of America

Subhashini Jagu

Affiliation Department of Pathology, The Johns Hopkins University, Baltimore, Maryland, United States of America

Reinhard Kirnbauer

Affiliation Laboratory of Viral Oncology, Division of Immunology, Allergy and Infectious Diseases, Department of Dermatology, Medical University Vienna (MUW), Vienna, Austria

Richard B. S. Roden

roden@jhmi.edu

Affiliations Department of Pathology, The Johns Hopkins University, Baltimore, Maryland, United States of America, Department of Gynecology and Obstetrics, The Johns Hopkins University, Baltimore, Maryland, United States of America, Department of Oncology, The Johns Hopkins University, Baltimore, Maryland, United States of America

Competing Interests

This study was partly funded by Sanofi Pasteur. Subhashini Jagu and Richard Roden are co-inventors on L2 patents (20090047301 Papillomavirus L2 N-Terminal Peptides for the Induction of Broadly Cross-Neutralizing Antibodies and 20100297144 MULTITYPE HPV PEPTIDE COMPOSITIONS AND METHODS FOR TREATMENT OR PREVENTION OF HUMAN PAPILLOMAVIRUS INFECTION), licensed to Shantha Biotechnics Ltd., GlaxoSmithKline, PaxVax, Inc., and Acambis, Inc., and have received research funding from Shantha Biotechnics Ltd., Sanofi Pasteur and GlaxoSmithKline. Richard Roden is a founder of Papivax LLC. The terms of these arrangements are managed by Johns Hopkins University in accordance with its conflict of interest policies. Reinhard Kirnbauer is a co-inventor on patent (7,361,356 Self-assembling recombinant papillomavirus capsid proteins), licensed to GlaxoSmithKline (GSK) and Merck (MSD). This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: RBSR KK. Performed the experiments: KK RJ JWW SJ. Analyzed the data: KK RJ JWW SJ RK RBSR. Contributed reagents/materials/analysis tools: RK RBSR. Wrote the paper: KK RK RBSR.