About the Authors

Kazuhiro Okumura

Affiliation Department of Carcinogenesis Research, Division of Experimental Animal Research, Chiba Cancer Center Research Institute, Chiba, Chiba, Japan

Megumi Saito

Affiliation Department of Carcinogenesis Research, Division of Experimental Animal Research, Chiba Cancer Center Research Institute, Chiba, Chiba, Japan

Eriko Isogai

Affiliation Department of Carcinogenesis Research, Division of Experimental Animal Research, Chiba Cancer Center Research Institute, Chiba, Chiba, Japan

Ikuo Miura

Affiliation Technology and Development Team for Mouse Phenotype Analysis, Japan Mouse Clinic, Riken Bioresource Center, Tsukuba, Ibaraki, Japan

Shigeharu Wakana

Affiliation Technology and Development Team for Mouse Phenotype Analysis, Japan Mouse Clinic, Riken Bioresource Center, Tsukuba, Ibaraki, Japan

Ryo Kominami

Affiliation Department of Molecular Genetics, Graduate School of Medical and Dental Sciences, Niigata University, Niigata, Niigata, Japan

Yuichi Wakabayashi

yuichi_wakabayashi@chiba-cc.jp

Affiliation Department of Carcinogenesis Research, Division of Experimental Animal Research, Chiba Cancer Center Research Institute, Chiba, Chiba, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KO YW. Performed the experiments: KO MS EI. Analyzed the data: IM SW. Contributed reagents/materials/analysis tools: RK. Contributed to the writing of the manuscript: KO YW.