About the Authors

Sawako Moriwaki

Contributed equally to this work with: Sawako Moriwaki, Keiko Suzuki

Affiliations Laboratory of Genomics and Proteomics, National Center for Geriatrics and Gerontology (NCGG), Aichi, Japan, Biobank Omics Unit, National Center for Geriatrics and Gerontology (NCGG), Aichi, Japan

Keiko Suzuki

Contributed equally to this work with: Sawako Moriwaki, Keiko Suzuki

Affiliation Department of Pharmacology, School of Dentistry, Showa University, Tokyo, Japan

Masashi Muramatsu

Current address: Department of Cancer Genetics, Roswell Park Cancer Institute, Elm and Carlton Streets, Buffalo, New York, United States of America

Affiliation Laboratory of Genomics and Proteomics, National Center for Geriatrics and Gerontology (NCGG), Aichi, Japan

Atsushi Nomura

Affiliation Nihon Seiyaku Kogyo, Co., Ltd., Aichi, Japan

Fumihide Inoue

Affiliation Nihon Seiyaku Kogyo, Co., Ltd., Aichi, Japan

Takeshi Into

Affiliation Department of Oral Bacteriology, Division of Oral Infections and Health Sciences, Asahi University School of Dentistry, Gifu, Japan

Yuji Yoshiko

Affiliation Department of Oral Growth and Developmental Biology, Hiroshima University Graduate School of Biomedical Sciences, Hiroshima, Japan

Shumpei Niida

sniida@ncgg.go.jp

Affiliations Laboratory of Genomics and Proteomics, National Center for Geriatrics and Gerontology (NCGG), Aichi, Japan, Biobank Omics Unit, National Center for Geriatrics and Gerontology (NCGG), Aichi, Japan

Competing Interests

AN and FI are employees of Nihon Seiyaku Kogyo Co., Ltd. Tama Biochemical Co. Ltd., who donated experimental material Cassis-extract-35, also declared that no competing interests exist. This does not alter adherence to all PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: SN. Performed the experiments: SM KS MM YY. Analyzed the data: KS TI YY. Contributed reagents/materials/analysis tools: AN FI. Wrote the paper: SN.