About the Authors

Karen Tse

Contributed equally to this work with: Karen Tse, Sreekanth Puttachary

Affiliation Institute of Ageing and Chronic Disease, University of Liverpool, Liverpool, United Kingdom

Sreekanth Puttachary

Contributed equally to this work with: Karen Tse, Sreekanth Puttachary

Affiliation Department of Biomedical Science, College of Veterinary Medicine, Iowa State University, Ames, Iowa, United States of America

Edward Beamer

Affiliation Institute of Ageing and Chronic Disease, University of Liverpool, Liverpool, United Kingdom

Graeme J. Sills

Affiliation Institute of Translational Medicine, University of Liverpool, Liverpool, United Kingdom

Thimmasettappa Thippeswamy

tswamy@iastate.edu

Affiliation Department of Biomedical Science, College of Veterinary Medicine, Iowa State University, Ames, Iowa, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TT. Performed the experiments: KT SP EB. Analyzed the data: KT SP EB. Wrote the paper: TT SP. Secured funding: TT. Recruited and trained personnel: TT. Proofread the first and the second drafts of the manuscript: GS.