About the Authors

Hashem Koohy

hashem.koohy@babraham.ac.uk (HK); th@sanger.ac.uk (TH)

Affiliations The Babraham Institute, Babraham Research Campus, Cambridge, United Kingdom, Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Cambridge, United Kingdom

Thomas A. Down

Affiliation The Babraham Institute, Babraham Research Campus, Cambridge, United Kingdom

Mikhail Spivakov

Affiliation Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Cambridge, United Kingdom

Tim Hubbard

hashem.koohy@babraham.ac.uk (HK); th@sanger.ac.uk (TH)

Affiliation The Babraham Institute, Babraham Research Campus, Cambridge, United Kingdom

Competing Interests

Timothy Hubbard is a member of PLOS ONE Editorial Board, however, this does not alter the author's adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: HK TH. Performed the experiments: HK TD MS. Analyzed the data: HK TD MS. Contributed reagents/materials/analysis tools: HK MS. Wrote the paper: HK TD MS TH.