About the Authors

Kasumi Ishida

Affiliation Department of Medical Laboratory Science, Faculty of Health Sciences, Hokkaido University, Sapporo, Hokkaido, Japan

Tsuyoshi Sekizuka

Affiliation Pathogen Genomics Center, National Institute of Infectious Diseases, Tokyo, Japan

Kyoko Hayashida

Affiliation Research Center for Zoonosis Control, Hokkaido University, Sapporo, Hokkaido, Japan

Junji Matsuo

Affiliation Department of Medical Laboratory Science, Faculty of Health Sciences, Hokkaido University, Sapporo, Hokkaido, Japan

Fumihiko Takeuchi

Affiliation Pathogen Genomics Center, National Institute of Infectious Diseases, Tokyo, Japan

Makoto Kuroda

Affiliation Pathogen Genomics Center, National Institute of Infectious Diseases, Tokyo, Japan

Shinji Nakamura

Affiliation Division of Biomedical Imaging Research, Juntendo University Graduate School of Medicine, Tokyo, Japan

Tomohiro Yamazaki

Affiliation Department of Medical Laboratory Science, Faculty of Health Sciences, Hokkaido University, Sapporo, Hokkaido, Japan

Mitsutaka Yoshida

Affiliation Division of Ultrastructural Research, Juntendo University Graduate School of Medicine, Tokyo, Japan

Kaori Takahashi

Affiliation Division of Ultrastructural Research, Juntendo University Graduate School of Medicine, Tokyo, Japan

Hiroki Nagai

Affiliation Research Institute for Microbial Diseases, Osaka University, Osaka, Japan

Chihiro Sugimoto

Affiliation Research Center for Zoonosis Control, Hokkaido University, Sapporo, Hokkaido, Japan

Hiroyuki Yamaguchi

hiroyuki@med.hokudai.ac.jp

Affiliation Department of Medical Laboratory Science, Faculty of Health Sciences, Hokkaido University, Sapporo, Hokkaido, Japan

Competing Interests

The authors declare that they have no competing interests.

Author Contributions

Conceived and designed the experiments: HY. Performed the experiments: JM TY KI SN MY KT. Analyzed the data: KI MK HY TS FT HN KH CS. Wrote the paper: HY.