About the Authors

Zbyszek Boratyński

boratyns@jyu.fi

Affiliations Division of Ecology and Evolutionary Biology, Department of Biological and Environmental Science, University of Jyväskylä, Jyväskylä, Finland, CIBIO/InBIO, Centro de Investigação em Biodiversidade e Recursos Genéticos, Universidade do Porto, Vairão, Portugal

José Carlos Brito

Affiliations CIBIO/InBIO, Centro de Investigação em Biodiversidade e Recursos Genéticos, Universidade do Porto, Vairão, Portugal, Departamento de Biologia, Faculdade de Ciências da Universidade do Porto, Porto, Portugal

João Carlos Campos

Affiliations CIBIO/InBIO, Centro de Investigação em Biodiversidade e Recursos Genéticos, Universidade do Porto, Vairão, Portugal, Departamento de Biologia, Faculdade de Ciências da Universidade do Porto, Porto, Portugal

Maija Karala

Affiliation Division of Ecology and Evolutionary Biology, Department of Biological and Environmental Science, University of Jyväskylä, Jyväskylä, Finland

Tapio Mappes

Affiliation Division of Ecology and Evolutionary Biology, Department of Biological and Environmental Science, University of Jyväskylä, Jyväskylä, Finland

Competing Interests

One of the authors, TM, is a PLOS ONE Editorial Board member. This does not alter adherence to PLOS ONE Editorial policies and criteria.

Author Contributions

Conceived and designed the experiments: ZB JC MK JCB. Performed the experiments: JC MK ZB. Analyzed the data: ZB JC MK TM JCB. Contributed reagents/materials/analysis tools: TM. Wrote the paper: ZB.