About the Authors

S. James Reynolds

J.Reynolds.2@bham.ac.uk (SJR); rasuk@btconnect.com (BJH)

Affiliations Centre for Ornithology, School of Biosciences, College of Life and Environmental Sciences, University of Birmingham, Edgbaston, Birmingham, United Kingdom, Army Ornithological Society, Land Forces Directorate, Army Headquarters, Andover, Hampshire, United Kingdom

Graham R. Martin

Affiliation Centre for Ornithology, School of Biosciences, College of Life and Environmental Sciences, University of Birmingham, Edgbaston, Birmingham, United Kingdom

Alistair Dawson

Affiliation Centre for Ecology and Hydrology, Penicuik, Midlothian, United Kingdom

Colin P. Wearn

Affiliation Royal Air Force Ornithological Society, Headquarters Air Command, Royal Air Force, High Wycombe, Buckinghamshire, United Kingdom

B. John Hughes

J.Reynolds.2@bham.ac.uk (SJR); rasuk@btconnect.com (BJH)

Affiliations Centre for Ornithology, School of Biosciences, College of Life and Environmental Sciences, University of Birmingham, Edgbaston, Birmingham, United Kingdom, Army Ornithological Society, Land Forces Directorate, Army Headquarters, Andover, Hampshire, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SJR GRM BJH. Performed the experiments: SJR CPW BJH. Analyzed the data: SJR AD BJH. Contributed reagents/materials/analysis tools: CPW. Wrote the paper: SJR GRM AD CPW BJH.