About the Authors

Joel A. Dietz

Contributed equally to this work with: Joel A. Dietz, Margaret E. Maes

Affiliation Department of Ophthalmology and Visual Sciences, University of Wisconsin, Madison, Wisconsin, United States of America

Margaret E. Maes

Contributed equally to this work with: Joel A. Dietz, Margaret E. Maes

Affiliation Department of Ophthalmology and Visual Sciences, University of Wisconsin, Madison, Wisconsin, United States of America

Shuang Huang

Affiliation Department of Biostatistics, University of Wisconsin, Madison, Wisconsin, United States of America

Brian S. Yandell

Affiliation Department of Biostatistics, University of Wisconsin, Madison, Wisconsin, United States of America

Cassandra L. Schlamp

Affiliation Department of Ophthalmology and Visual Sciences, University of Wisconsin, Madison, Wisconsin, United States of America

Angela D. Montgomery

Affiliation Department of Ophthalmology and Visual Sciences, University of Wisconsin, Madison, Wisconsin, United States of America

R. Rand Allingham

Affiliation Center for Human Genetics, Department of Medicine, Duke University Medical Center, Durham, North Carolina, United States of America

Michael A. Hauser

Affiliation Center for Human Genetics, Department of Medicine, Duke University Medical Center, Durham, North Carolina, United States of America

Robert W. Nickells

nickells@wisc.edu

Affiliation Department of Ophthalmology and Visual Sciences, University of Wisconsin, Madison, Wisconsin, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RWN BSY CLS MEM RRA MAH. Performed the experiments: JAD MEM ADM CLS RWN. Analyzed the data: SH BSY RWN JAD MEM CLS ADM. Wrote the paper: RWN CLS MEM JAD RRA MAH BSY.