About the Authors

Si Tan

Contributed equally to this work with: Si Tan, Mingxia Li

Affiliations College of Horticulture and Landscape Architecture, Southwest University, Chongqing, China, Drug Discovery Lab, School of Pharmacy, Shanghai University of Traditional Chinese Medicine, Shanghai, China, Key Laboratory of Horticulture Science for Southern Mountainous Regions, Ministry of Education, Chongqing, China

Mingxia Li

Contributed equally to this work with: Si Tan, Mingxia Li

Affiliations College of Horticulture and Landscape Architecture, Southwest University, Chongqing, China, Drug Discovery Lab, School of Pharmacy, Shanghai University of Traditional Chinese Medicine, Shanghai, China, Key Laboratory of Horticulture Science for Southern Mountainous Regions, Ministry of Education, Chongqing, China

Xiaobo Ding

Affiliations College of Horticulture and Landscape Architecture, Southwest University, Chongqing, China, Drug Discovery Lab, School of Pharmacy, Shanghai University of Traditional Chinese Medicine, Shanghai, China, Key Laboratory of Horticulture Science for Southern Mountainous Regions, Ministry of Education, Chongqing, China

Shengjie Fan

Affiliation Drug Discovery Lab, School of Pharmacy, Shanghai University of Traditional Chinese Medicine, Shanghai, China

Lu Guo

Affiliation Drug Discovery Lab, School of Pharmacy, Shanghai University of Traditional Chinese Medicine, Shanghai, China

Ming Gu

Affiliation Drug Discovery Lab, School of Pharmacy, Shanghai University of Traditional Chinese Medicine, Shanghai, China

Yu Zhang

Affiliation Drug Discovery Lab, School of Pharmacy, Shanghai University of Traditional Chinese Medicine, Shanghai, China

Li Feng

Affiliation Drug Discovery Lab, School of Pharmacy, Shanghai University of Traditional Chinese Medicine, Shanghai, China

Dong Jiang

Affiliation Citrus Research Institute, Chinese Academy of Agricultural Sciences, Chongqing, China

Yiming Li

Affiliation Drug Discovery Lab, School of Pharmacy, Shanghai University of Traditional Chinese Medicine, Shanghai, China

Wanpeng Xi

Affiliations College of Horticulture and Landscape Architecture, Southwest University, Chongqing, China, Key Laboratory of Horticulture Science for Southern Mountainous Regions, Ministry of Education, Chongqing, China

Cheng Huang

zzqswu@yahoo.com (ZZ); chuang.shutcm@gmail.com (CH)

Affiliation Drug Discovery Lab, School of Pharmacy, Shanghai University of Traditional Chinese Medicine, Shanghai, China

Zhiqin Zhou

zzqswu@yahoo.com (ZZ); chuang.shutcm@gmail.com (CH)

Affiliations College of Horticulture and Landscape Architecture, Southwest University, Chongqing, China, Key Laboratory of Horticulture Science for Southern Mountainous Regions, Ministry of Education, Chongqing, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CH ZZ. Performed the experiments: ST ML XD SF LG MG YZ LF. Analyzed the data: ST ML XD WX CH. Contributed reagents/materials/analysis tools: DJ YL. Wrote the paper: ST ML WX ZZ.