About the Authors

Naoki Katayama

katayama6@affrc.go.jp

Affiliation Biodiversity Division, National Institute for Agro-Environmental Sciences, Tsukuba-shi, Ibaraki, Japan

Tatsuya Amano

Affiliation Conservation Science Group, Department of Zoology, University of Cambridge, Cambridge, United Kingdom

Shoji Naoe

Affiliation Laboratory of Biodiversity Science, School of Agricultural and Life Sciences, University of Tokyo, Tokyo, Japan

Takehisa Yamakita

Affiliation Japan Agency for Marine-Earth Science and Technology (JAMSTEC), Yokosuka, Kanagawa, Japan

Isamu Komatsu

Affiliation Laboratory of Biodiversity Science, School of Agricultural and Life Sciences, University of Tokyo, Tokyo, Japan

Shin-ichi Takagawa

Affiliation The Nature Conservation Society of Japan, Chuo-ku, Tokyo, Japan

Naoto Sato

Affiliation Biodiversity Center of Japan, Kamiyoshida, Fujiyoshida-shi, Yamanashi, Japan

Mutsuyuki Ueta

Affiliation Japan Bird Research Association, Fuchu-shi, Tokyo, Japan

Tadashi Miyashita

Affiliation Laboratory of Biodiversity Science, School of Agricultural and Life Sciences, University of Tokyo, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ST NS MU TM. Analyzed the data: NK TA SN TY IK. Contributed reagents/materials/analysis tools: SN TY ST MU. Wrote the paper: NK TA TM. Obtained permission for the use of bird data: ST NS MU.