About the Authors

Hiroshi Saga

saga@ono.co.jp

Affiliation Exploratory Research Laboratories, ONO Pharmaceutical Co., Ltd., Tsukuba, Ibaraki, Japan

Akira Ohhata

Affiliation Medicinal Chemistry Research Laboratories, ONO Pharmaceutical Co., Ltd., Shimamoto, Mishima, Osaka, Japan

Akio Hayashi

Affiliation Exploratory Research Laboratories, ONO Pharmaceutical Co., Ltd., Tsukuba, Ibaraki, Japan

Makoto Katoh

Affiliation Exploratory Research Laboratories, ONO Pharmaceutical Co., Ltd., Tsukuba, Ibaraki, Japan

Tatsuo Maeda

Affiliation Exploratory Research Laboratories, ONO Pharmaceutical Co., Ltd., Tsukuba, Ibaraki, Japan

Hirotaka Mizuno

Affiliation Exploratory Research Laboratories, ONO Pharmaceutical Co., Ltd., Tsukuba, Ibaraki, Japan

Yuka Takada

Affiliation Exploratory Research Laboratories, ONO Pharmaceutical Co., Ltd., Tsukuba, Ibaraki, Japan

Yuka Komichi

Affiliation Exploratory Research Laboratories, ONO Pharmaceutical Co., Ltd., Tsukuba, Ibaraki, Japan

Hiroto Ota

Affiliation Exploratory Research Laboratories, ONO Pharmaceutical Co., Ltd., Tsukuba, Ibaraki, Japan

Naoya Matsumura

Affiliation Discovery Technology Laboratories, ONO Pharmaceutical Co., Ltd., Shimamoto, Mishima, Osaka, Japan

Masami Shibaya

Affiliation Safety Research Laboratories, ONO Phamaceutical Co., Ltd., Sakai, Fukui, Japan

Tetsuya Sugiyama

Affiliation Exploratory Research Laboratories, ONO Pharmaceutical Co., Ltd., Tsukuba, Ibaraki, Japan

Shinji Nakade

Affiliation Exploratory Research Laboratories, ONO Pharmaceutical Co., Ltd., Tsukuba, Ibaraki, Japan

Katsuya Kishikawa

Affiliation Exploratory Research Laboratories, ONO Pharmaceutical Co., Ltd., Tsukuba, Ibaraki, Japan

Competing Interests

All authors are employees of Ono Pharmaceutical Co. Ltd. This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: HS TS SN KK. Performed the experiments: HS AO AH MK TM HM YT YK HO NM MS. Analyzed the data: HS AH MK TM YT YK NM MS. Contributed reagents/materials/analysis tools: HS AO MK TM TS. Wrote the paper: HS AH MK TM KK.