About the Authors

Neil C. Sheppard

Affiliation Vaccine Research, Worldwide R&D, Pfizer Inc., San Diego, California, United States of America

R. Brad Jones

Affiliation Department of Immunology, University of Toronto, Toronto, ON, Canada

Benjamin J. Burwitz

Affiliations Vaccine and Gene Therapy Institute, Oregon Health & Science University, Beaverton, Oregon, United States of America, Oregon National Primate Research Center, Oregon Health & Science University, Beaverton, Oregon, United States of America

Francesca A. Nimityongskul

Affiliation Wisconsin National Primate Center, University of Wisconsin, Madison, Wisconsin, United States of America

Laura P. Newman

Affiliation Wisconsin National Primate Center, University of Wisconsin, Madison, Wisconsin, United States of America

Matthew B. Buechler

Affiliation Wisconsin National Primate Center, University of Wisconsin, Madison, Wisconsin, United States of America

Jason S. Reed

Affiliations Vaccine and Gene Therapy Institute, Oregon Health & Science University, Beaverton, Oregon, United States of America, Oregon National Primate Research Center, Oregon Health & Science University, Beaverton, Oregon, United States of America

Shari M. Piaskowski

Affiliation Wisconsin National Primate Center, University of Wisconsin, Madison, Wisconsin, United States of America

Kim L. Weisgrau

Affiliation Wisconsin National Primate Center, University of Wisconsin, Madison, Wisconsin, United States of America

Philip A. Castrovinci

Affiliation Wisconsin National Primate Center, University of Wisconsin, Madison, Wisconsin, United States of America

Nancy A. Wilson

Affiliation Wisconsin National Primate Center, University of Wisconsin, Madison, Wisconsin, United States of America

Mario A. Ostrowski

Affiliation Department of Immunology, University of Toronto, Toronto, ON, Canada

Byung Park

Affiliation Division of Biostatistics, Department of Public Health and Preventive Medicine, Oregon Health and Science University, Portland, Oregon, United States of America

Douglas F. Nixon

Affiliation Division of Experiment Medicine, San Francisco General Hospital, University of California San Francisco, San Francisco, California, United States of America

Eva G. Rakasz

Affiliation Wisconsin National Primate Center, University of Wisconsin, Madison, Wisconsin, United States of America

Jonah B. Sacha

sacha@ohsu.edu

Affiliations Vaccine and Gene Therapy Institute, Oregon Health & Science University, Beaverton, Oregon, United States of America, Oregon National Primate Research Center, Oregon Health & Science University, Beaverton, Oregon, United States of America, Department of Medical Microbiology and Immunology, Oregon Health & Science University, Portland, Oregon, United States of America

Competing Interests

Neil Sheppard was previously employed by Vaccine Research, Worldwide R&D, Pfizer Inc., San Diego, CA, USA. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: NCS RBJ MAO DFN JBS. Performed the experiments: NCS RBJ FAN LPN MBB JSR SMP KLW PAC EGR. Analyzed the data: NCS RBJ BJB JSR BP JBS. Contributed reagents/materials/analysis tools: NAW RBJ MAO. Wrote the paper: NCS RBJ BJB JBS.