About the Authors

Xing Ma

Affiliations Stowers Institute for Medical Research, Kansas City, Missouri, United States of America, Department of Anatomy and Cell Biology, University of Kansas School of Medicine, Kansas City, Kansas, United States of America

Su Wang

Affiliations Stowers Institute for Medical Research, Kansas City, Missouri, United States of America, Department of Anatomy and Cell Biology, University of Kansas School of Medicine, Kansas City, Kansas, United States of America

Trieu Do

Affiliation Stowers Institute for Medical Research, Kansas City, Missouri, United States of America

Xiaoqing Song

Affiliation Stowers Institute for Medical Research, Kansas City, Missouri, United States of America

Mayu Inaba

Affiliation Life Sciences Institute, Center for Stem Cell Biology, University of Michigan, Ann Arbor, Michigan, United States of America

Yoshiya Nishimoto

Affiliation Stowers Institute for Medical Research, Kansas City, Missouri, United States of America

Lu-ping Liu

Affiliation School of Medicine, Tsinghua University, Beijing, China

Yuan Gao

Affiliation School of Medicine, Tsinghua University, Beijing, China

Ying Mao

Affiliation School of Medicine, Tsinghua University, Beijing, China

Hui Li

Affiliation School of Medicine, Tsinghua University, Beijing, China

William McDowell

Affiliation Stowers Institute for Medical Research, Kansas City, Missouri, United States of America

Jungeun Park

Affiliation Stowers Institute for Medical Research, Kansas City, Missouri, United States of America

Kate Malanowski

Affiliation Stowers Institute for Medical Research, Kansas City, Missouri, United States of America

Allison Peak

Affiliation Stowers Institute for Medical Research, Kansas City, Missouri, United States of America

Anoja Perera

Affiliation Stowers Institute for Medical Research, Kansas City, Missouri, United States of America

Hua Li

Affiliation Stowers Institute for Medical Research, Kansas City, Missouri, United States of America

Karin Gaudenz

Affiliation Stowers Institute for Medical Research, Kansas City, Missouri, United States of America

Jeff Haug

Affiliation Stowers Institute for Medical Research, Kansas City, Missouri, United States of America

Yukiko Yamashita

Affiliation Life Sciences Institute, Center for Stem Cell Biology, University of Michigan, Ann Arbor, Michigan, United States of America

Haifan Lin

Affiliation Yale Stem Cell Center, Yale University School of Medicine, New Haven, Connecticut, United Sates of America

Jian-quan Ni

Affiliation School of Medicine, Tsinghua University, Beijing, China

Ting Xie

tgx@stowers.org

Affiliations Stowers Institute for Medical Research, Kansas City, Missouri, United States of America, Department of Anatomy and Cell Biology, University of Kansas School of Medicine, Kansas City, Kansas, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: XM TX. Performed the experiments: XM SW TD XS YN LL YG YM Hui Li KM A. Peak A. Perera Hua Li WM JP KG JH JN. Analyzed the data: Hua Li XM TX. Contributed reagents/materials/analysis tools: MI YY Haifan Lin. Wrote the paper: XM TX.