About the Authors

Peter J. Unmack

peter.pub@unmack.net

Affiliations WIDB 401, Department of Biology, Brigham Young University, Provo, Utah, United States of America, National Evolutionary Synthesis Center, Durham, North Carolina, United States of America

Thomas E. Dowling

Affiliations School of Life Sciences, Arizona State University, Tempe, Arizona, United States of America, Department of Biological Sciences, Wayne State University, Detroit, Michigan, United States of America

Nina J. Laitinen

Affiliation WIDB 401, Department of Biology, Brigham Young University, Provo, Utah, United States of America

Carol L. Secor

Affiliation School of Life Sciences, Arizona State University, Tempe, Arizona, United States of America

Richard L. Mayden

Affiliation Department of Biology, Saint Louis University, St. Louis, Missouri, United States of America

Dennis K. Shiozawa

Affiliation WIDB 401, Department of Biology, Brigham Young University, Provo, Utah, United States of America

Gerald R. Smith

Affiliation Museum of Zoology, The University of Michigan, Ann Arbor, Michigan, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: PJU TED NJL CLS RLM DKS GRS. Performed the experiments: PJU TED NJL CLS RLM GRS. Analyzed the data: PJU TED GRS. Contributed reagents/materials/analysis tools: PJU TED NJL CLS RLM DKS GRS. Wrote the paper: PJU TED NJL CLS RLM DKS GRS.