About the Authors

Hee-Soo Park

Affiliation Department of Bacteriology, University of Wisconsin, Madison, Wisconsin, United States of America

Tae-Young Nam

Affiliation Department of Bacteriology, University of Wisconsin, Madison, Wisconsin, United States of America

Kap-Hoon Han

Affiliations Department of Bacteriology, University of Wisconsin, Madison, Wisconsin, United States of America, Department of Pharmaceutical Engineering, Woosuk University, Wanju, Republic of Korea

Sun Chang Kim

Affiliation Department of Biological Sciences, Korea Advanced Institute of Science and Technology, Dae-Jon, Republic of Korea

Jae-Hyuk Yu

jyu1@wisc.edu

Affiliations Department of Bacteriology, University of Wisconsin, Madison, Wisconsin, United States of America, Department of Genetics, University of Wisconsin, Madison, Wisconsin, United States of America

Competing Interests

Jae-Hyuk Yu and Kap-Hoon Han are Academic Editors of PLoS ONE. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: H-SP S-CK J-HY. Performed the experiments: H-SP T-YN. Analyzed the data: H-SP T-YN K-HH. Contributed reagents/materials/analysis tools: K-HH S-CK J-HY. Wrote the paper: H-SP J-HY.