About the Authors

Kirsty I. Munro

Contributed equally to this work with: Kirsty I. Munro, Michael J. Thrippleton

Affiliation MRC Centre for Reproductive Health, University of Edinburgh, Edinburgh, United Kingdom

Michael J. Thrippleton

Contributed equally to this work with: Kirsty I. Munro, Michael J. Thrippleton

Affiliations MRC Centre for Reproductive Health, University of Edinburgh, Edinburgh, United Kingdom, Centre for Clinical Brain Sciences, University of Edinburgh, Edinburgh, United Kingdom

Alistair R. W. Williams

Affiliations MRC Centre for Reproductive Health, University of Edinburgh, Edinburgh, United Kingdom, Royal Infirmary of Edinburgh, NHS Lothian, Edinburgh, United Kingdom

Graham McKillop

Affiliation Royal Infirmary of Edinburgh, NHS Lothian, Edinburgh, United Kingdom

Jane Walker

Affiliation Royal Infirmary of Edinburgh, NHS Lothian, Edinburgh, United Kingdom

Andrew W. Horne

Affiliation MRC Centre for Reproductive Health, University of Edinburgh, Edinburgh, United Kingdom

David E. Newby

Affiliation Centre for Cardiovascular Science, University of Edinburgh, Edinburgh, United Kingdom

Richard A. Anderson

Affiliation MRC Centre for Reproductive Health, University of Edinburgh, Edinburgh, United Kingdom

Scott I. Semple

Affiliations Centre for Cardiovascular Science, University of Edinburgh, Edinburgh, United Kingdom, Clinical Research Imaging Centre, University of Edinburgh, Edinburgh, United Kingdom

Ian Marshall

Affiliations Centre for Clinical Brain Sciences, University of Edinburgh, Edinburgh, United Kingdom, Centre for Cardiovascular Science, University of Edinburgh, Edinburgh, United Kingdom

Steff C. Lewis

Affiliation Centre for Population Health Sciences, University of Edinburgh, Edinburgh, United Kingdom

Robert P. Millar

Affiliations MRC Centre for Reproductive Health, University of Edinburgh, Edinburgh, United Kingdom, Mammal Research Institute, University of Pretoria, Pretoria, South Africa, UCT/MRC Receptor Biology Unit, University of Cape Town, Cape Town, South Africa, Centre for Integrative Physiology, University of Edinburgh, Edinburgh, United Kingdom

Mark E. Bastin

Affiliation Centre for Clinical Brain Sciences, University of Edinburgh, Edinburgh, United Kingdom

Hilary O. D. Critchley

hilary.critchley@ed.ac.uk

Affiliation MRC Centre for Reproductive Health, University of Edinburgh, Edinburgh, United Kingdom

Competing Interests

HODC is a PLOS ONE Editorial Board member (since 2012 an Academic Editor). She confirms that this does not alter her adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: ARW GM JW AWH RAA SIS IM SL RPM MEB HODC. Performed the experiments: KIM MJT GM JW SIS IM MEB. Analyzed the data: KIM MJT ARW GM JW RAA SL. Wrote the paper: KIM MJT HODC DEN ARW GM JW AWH RAA SIS IM SL RPM MEB.