About the Authors

Jennifer Olt

Contributed equally to this work with: Jennifer Olt, Philomena Mburu

Affiliation Department of Biomedical Science, University of Sheffield, Sheffield, United Kingdom

Philomena Mburu

Contributed equally to this work with: Jennifer Olt, Philomena Mburu

Affiliation Medical Research Council (MRC), Mammalian Genetics Unit, Harwell, United Kingdom

Stuart L. Johnson

Affiliation Department of Biomedical Science, University of Sheffield, Sheffield, United Kingdom

Andy Parker

Affiliation Medical Research Council (MRC), Mammalian Genetics Unit, Harwell, United Kingdom

Stephanie Kuhn

Current address: Department of Biophysics, Saarland University, Homburg, Germany

Affiliation Department of Biomedical Science, University of Sheffield, Sheffield, United Kingdom

Mike Bowl

Affiliation Medical Research Council (MRC), Mammalian Genetics Unit, Harwell, United Kingdom

Walter Marcotti

w.marcotti@sheffield.ac.uk (WM); S.Brown@har.mrc.ac.uk (SDMB)

Affiliation Department of Biomedical Science, University of Sheffield, Sheffield, United Kingdom

Steve D. M. Brown

w.marcotti@sheffield.ac.uk (WM); S.Brown@har.mrc.ac.uk (SDMB)

Affiliation Medical Research Council (MRC), Mammalian Genetics Unit, Harwell, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: WM SDMB. Performed the experiments: JO PM SLJ AP SK MB WM SDMB. Analyzed the data: JO PM SLJ AP SK MB WM SDMB. Wrote the paper: WM SDMB.