About the Authors

Masato Kiyoshi

Affiliation Medical Genome Sciences, Graduate School of Frontier Sciences, The University of Tokyo, Kashiwa, Chiba, Japan

Jose M. M. Caaveiro

Affiliations Department of Bioengineering, Graduate School of Engineering, The University of Tokyo, Bunkyo-ku, Tokyo, Japan, Laboratory of Medical Proteomics, Institute of Medical Science, The University of Tokyo, Minato-ku, Tokyo, Japan

Eri Miura

Affiliation Medical Genome Sciences, Graduate School of Frontier Sciences, The University of Tokyo, Kashiwa, Chiba, Japan

Satoru Nagatoishi

Affiliation Department of Bioengineering, Graduate School of Engineering, The University of Tokyo, Bunkyo-ku, Tokyo, Japan

Makoto Nakakido

Affiliation Laboratory of Medical Proteomics, Institute of Medical Science, The University of Tokyo, Minato-ku, Tokyo, Japan

Shinji Soga

Affiliation Molecular Medicine Research Laboratories, Drug Discovery Research, Astellas Pharma Inc., Tsukuba, Ibaraki, Japan

Hiroki Shirai

Affiliation Molecular Medicine Research Laboratories, Drug Discovery Research, Astellas Pharma Inc., Tsukuba, Ibaraki, Japan

Shigeki Kawabata

Affiliation Molecular Medicine Research Laboratories, Drug Discovery Research, Astellas Pharma Inc., Tsukuba, Ibaraki, Japan

Kouhei Tsumoto

tsumoto@bioeng.t.u-tokyo.ac.jp

Affiliations Medical Genome Sciences, Graduate School of Frontier Sciences, The University of Tokyo, Kashiwa, Chiba, Japan, Department of Bioengineering, Graduate School of Engineering, The University of Tokyo, Bunkyo-ku, Tokyo, Japan, Laboratory of Medical Proteomics, Institute of Medical Science, The University of Tokyo, Minato-ku, Tokyo, Japan, Department of Chemistry and Biotechnology, Graduate School of Engineering, The University of Tokyo, Bunkyo-ku, Tokyo, Japan

Competing Interests

SS, HS, and SK are full-time employees of Astellas Pharma Inc. KT serves as a consultant for Astellas Pharma Inc. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: MK MN SS HS SK KT. Performed the experiments: MK EM MN SS. Analyzed the data: MK JMMC SN SS HS SK KT. Contributed reagents/materials/analysis tools: MK EM MN SS HS SK KT. Wrote the paper: MK JMMC SN HS KT.