About the Authors

Barbara B. R. Raddatz

Contributed equally to this work with: Barbara B. R. Raddatz, Florian Hansmann, Ingo Spitzbarth

Affiliations Department of Pathology, University of Veterinary Medicine Hannover, Hannover, Germany, Center for Systems Neuroscience, University of Veterinary Medicine Hannover, Hannover, Germany

Florian Hansmann

Contributed equally to this work with: Barbara B. R. Raddatz, Florian Hansmann, Ingo Spitzbarth

Affiliations Department of Pathology, University of Veterinary Medicine Hannover, Hannover, Germany, Center for Systems Neuroscience, University of Veterinary Medicine Hannover, Hannover, Germany

Ingo Spitzbarth

Contributed equally to this work with: Barbara B. R. Raddatz, Florian Hansmann, Ingo Spitzbarth

Affiliations Department of Pathology, University of Veterinary Medicine Hannover, Hannover, Germany, Center for Systems Neuroscience, University of Veterinary Medicine Hannover, Hannover, Germany

Arno Kalkuhl

Affiliation Department of Non-Clinical Drug Safety, Boehringer Ingelheim Pharma GmbH&Co KG, Biberach (Riß), Germany

Ulrich Deschl

Affiliation Department of Non-Clinical Drug Safety, Boehringer Ingelheim Pharma GmbH&Co KG, Biberach (Riß), Germany

Wolfgang Baumgärtner

Affiliations Department of Pathology, University of Veterinary Medicine Hannover, Hannover, Germany, Center for Systems Neuroscience, University of Veterinary Medicine Hannover, Hannover, Germany

Reiner Ulrich

Reiner.Ulrich@tiho-hannover.de

Affiliations Department of Pathology, University of Veterinary Medicine Hannover, Hannover, Germany, Center for Systems Neuroscience, University of Veterinary Medicine Hannover, Hannover, Germany

Competing Interests

Arno Kalkuhl and Ulrich Deschl are employed by Boehringer Ingelheim Pharma GmbH&Co KG. There are no patents, products in development or marketed products to declare. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: RU BR FH IS AK UD WB. Analyzed the data: RU BR FH IS. Contributed reagents/materials/analysis tools: RU BR FH IS AK UD WB. Wrote the paper: RU BR FH IS WB.