About the Authors

Jamie C. Prentice

Affiliations Disease Systems Team, SRUC, Edinburgh, United Kingdom, Biomathematics and Statistics Scotland, Edinburgh, United Kingdom, Environment Department, University of York, York, United Kingdom

Glenn Marion

glenn@bioss.ac.uk

Affiliation Biomathematics and Statistics Scotland, Edinburgh, United Kingdom

Piran C. L. White

Affiliation Environment Department, University of York, York, United Kingdom

Ross S. Davidson

Affiliation Disease Systems Team, SRUC, Edinburgh, United Kingdom

Michael R. Hutchings

Affiliation Disease Systems Team, SRUC, Edinburgh, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JCP GM RSD PCLW MRH. Performed the experiments: JCP GM RSD MRH. Analyzed the data: JCP GM RSD MRH. Wrote the paper: JCP GM RSD PCLW MRH.