About the Authors

Jot Hui Ooi

Affiliation Center for Molecular Immunology and Infectious Disease, Department of Biochemistry and Molecular Biology, Department of Veterinary and Biomedical Science, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Amanda Waddell

Affiliation Center for Molecular Immunology and Infectious Disease, Department of Biochemistry and Molecular Biology, Department of Veterinary and Biomedical Science, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Yang-Ding Lin

Affiliation Center for Molecular Immunology and Infectious Disease, Department of Biochemistry and Molecular Biology, Department of Veterinary and Biomedical Science, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Istvan Albert

Affiliation Center for Molecular Immunology and Infectious Disease, Department of Biochemistry and Molecular Biology, Department of Veterinary and Biomedical Science, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Laura T. Rust

Affiliation Center for Molecular Immunology and Infectious Disease, Department of Biochemistry and Molecular Biology, Department of Veterinary and Biomedical Science, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Victoria Holden

Affiliation Center for Molecular Immunology and Infectious Disease, Department of Biochemistry and Molecular Biology, Department of Veterinary and Biomedical Science, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Margherita T. Cantorna

mxc69@psu.edu

Affiliation Center for Molecular Immunology and Infectious Disease, Department of Biochemistry and Molecular Biology, Department of Veterinary and Biomedical Science, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JHO MTC. Performed the experiments: JHO AW YL LTR VH. Analyzed the data: JHO AW YL IA LTR VH MTC. Wrote the paper: JHO MTC.