About the Authors

Yan Sun

Contributed equally to this work with: Yan Sun, Rui Hou

Affiliation Key Laboratory of Marine Genetics and Breeding (MGB), Ministry of Education, College of Marine Life Sciences, Ocean University of China, Qingdao, China

Rui Hou

Contributed equally to this work with: Yan Sun, Rui Hou

Affiliation Key Laboratory of Marine Genetics and Breeding (MGB), Ministry of Education, College of Marine Life Sciences, Ocean University of China, Qingdao, China

Xiaoteng Fu

Affiliation Key Laboratory of Marine Genetics and Breeding (MGB), Ministry of Education, College of Marine Life Sciences, Ocean University of China, Qingdao, China

Changsen Sun

Affiliation School of Life Science, Taizhou University, Taizhou, China

Shi Wang

Affiliation Key Laboratory of Marine Genetics and Breeding (MGB), Ministry of Education, College of Marine Life Sciences, Ocean University of China, Qingdao, China

Chen Wang

Affiliation Key Laboratory of Marine Genetics and Breeding (MGB), Ministry of Education, College of Marine Life Sciences, Ocean University of China, Qingdao, China

Ning Li

Affiliation Key Laboratory of Marine Genetics and Breeding (MGB), Ministry of Education, College of Marine Life Sciences, Ocean University of China, Qingdao, China

Lingling Zhang

lingling80@ouc.edu.cn

Affiliation Key Laboratory of Marine Genetics and Breeding (MGB), Ministry of Education, College of Marine Life Sciences, Ocean University of China, Qingdao, China

Zhenmin Bao

Affiliation Key Laboratory of Marine Genetics and Breeding (MGB), Ministry of Education, College of Marine Life Sciences, Ocean University of China, Qingdao, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ZB LZ. Performed the experiments: YS RH XF CW NL. Analyzed the data: YS RH XF SW LZ. Wrote the paper: YS RH XF CS SW LZ ZB.