About the Authors

Mi Jin Kim

Contributed equally to this work with: Mi Jin Kim, Hoon Jae Jeong

Affiliation Department of Biomedical Science and Research Institute of Bioscience and Biotechnology, Hallym University, Chunchon, Gangwondo, Korea

Hoon Jae Jeong

Contributed equally to this work with: Mi Jin Kim, Hoon Jae Jeong

Affiliation Department of Biomedical Science and Research Institute of Bioscience and Biotechnology, Hallym University, Chunchon, Gangwondo, Korea

Dae Won Kim

Affiliation Department of Biochemistry and Molecular Biology, Research Institute of Oral Sciences, College of Dentistry, Kangnung-Wonju National University, Gangneung, Gangwondo, Korea

Eun Jeong Sohn

Affiliation Department of Biomedical Science and Research Institute of Bioscience and Biotechnology, Hallym University, Chunchon, Gangwondo, Korea

Hyo Sang Jo

Affiliation Department of Biomedical Science and Research Institute of Bioscience and Biotechnology, Hallym University, Chunchon, Gangwondo, Korea

Duk-Soo Kim

Affiliation Department of Anatomy, College of Medicine, Soonchunhyang University, Cheonan-Si, Chungcheonnamdo, Korea

Hyun Ah Kim

Affiliation Division of Rheumatology, Department of Internal Medicine, Hallym University Sacred Heart Hospital, Pyongchon, Kyunggido, Korea

Eun Young Park

Affiliation Department of Biological Sciences, Sookmyung Women’s University, Seoul, Korea

Jong Hoon Park

Affiliation Department of Biological Sciences, Sookmyung Women’s University, Seoul, Korea

Ora Son

Affiliation Department of Biomedical Science and Research Institute of Bioscience and Biotechnology, Hallym University, Chunchon, Gangwondo, Korea

Kyu Hyung Han

Affiliation Department of Biomedical Science and Research Institute of Bioscience and Biotechnology, Hallym University, Chunchon, Gangwondo, Korea

Jinseu Park

Affiliation Department of Biomedical Science and Research Institute of Bioscience and Biotechnology, Hallym University, Chunchon, Gangwondo, Korea

Won Sik Eum

wseum@hallym.ac.kr (WSE); sychoi@hallym.ac.kr (SYC)

Affiliation Department of Biomedical Science and Research Institute of Bioscience and Biotechnology, Hallym University, Chunchon, Gangwondo, Korea

Soo Young Choi

wseum@hallym.ac.kr (WSE); sychoi@hallym.ac.kr (SYC)

Affiliation Department of Biomedical Science and Research Institute of Bioscience and Biotechnology, Hallym University, Chunchon, Gangwondo, Korea

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: WSE SYC. Performed the experiments: MJK HJJ DWK EJS HSJ DSK. Analyzed the data: HAK JHP KHH JP WSE SYC. Contributed reagents/materials/analysis tools: EYP OS. Wrote the paper: WSE SYC.