About the Authors

Toshiyuki Mizui

Affiliations Department of Neurobiology and Behavior, Gunma University Graduate School of Medicine, Maebashi, Gunma, Japan, Bio-interface Research Group, Health Research Institute, National Institute of Advanced Industrial Science and Technology (AIST), Ikeda, Osaka, Japan

Yuko Sekino

yukos@nihs.go.jp (YS); tshirao@ gunma-u.ac.jp (TS)

Current address: Division of Pharmacology, National Institute of Health Sciences, Setagaya-ku, Tokyo, Japan

Affiliations Department of Neurobiology and Behavior, Gunma University Graduate School of Medicine, Maebashi, Gunma, Japan, Core Research for Evolution Science and Technology, Japan Science and Technology Corporation, Kawaguchi, Saitama, Japan

Hiroyuki Yamazaki

Affiliation Department of Neurobiology and Behavior, Gunma University Graduate School of Medicine, Maebashi, Gunma, Japan

Yuta Ishizuka

Affiliation Department of Neurobiology and Behavior, Gunma University Graduate School of Medicine, Maebashi, Gunma, Japan

Hideto Takahashi

Current address: Institut de recherches cliniques de Montréal (IRCM), Université de Montréal (UdeM), Québec, Canada

Affiliation Department of Neurobiology and Behavior, Gunma University Graduate School of Medicine, Maebashi, Gunma, Japan

Nobuhiko Kojima

Affiliation Department of Neurobiology and Behavior, Gunma University Graduate School of Medicine, Maebashi, Gunma, Japan

Masami Kojima

Affiliations Core Research for Evolution Science and Technology, Japan Science and Technology Corporation, Kawaguchi, Saitama, Japan, Bio-interface Research Group, Health Research Institute, National Institute of Advanced Industrial Science and Technology (AIST), Ikeda, Osaka, Japan

Tomoaki Shirao

yukos@nihs.go.jp (YS); tshirao@ gunma-u.ac.jp (TS)

Affiliation Department of Neurobiology and Behavior, Gunma University Graduate School of Medicine, Maebashi, Gunma, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TM YS TS. Performed the experiments: TM HY YI HT. Analyzed the data: TM YS HY YI HT NK MK. Wrote the paper: TM YS TS.