About the Authors

Emilie Donadieu

Affiliations Virology (UMR1161), French National Institute for Agricultural Research (INRA), Maisons-Alfort, France, Virology (UMR1161), French Agency for Food, Environmental and Occupational Health and Safety (ANSES), Maisons-Alfort, France, Virology (UMR1161), Paris-Est University, National Veterinary School of Alfort, Maisons-Alfort, France

Steeve Lowenski

Affiliations Virology (UMR1161), French National Institute for Agricultural Research (INRA), Maisons-Alfort, France, Virology (UMR1161), French Agency for Food, Environmental and Occupational Health and Safety (ANSES), Maisons-Alfort, France, Virology (UMR1161), Paris-Est University, National Veterinary School of Alfort, Maisons-Alfort, France

Jean-Luc Servely

Affiliations French National Institute for Agricultural Research (INRA), Nouzilly, France, Histology and Pathological Anatomy, Paris-Est University, National Veterinary School of Alfort, Maisons-Alfort, France

Eve Laloy

Affiliation Histology and Pathological Anatomy, Paris-Est University, National Veterinary School of Alfort, Maisons-Alfort, France

Thomas Lilin

Affiliation Biomedical Research Center, Paris-Est University, National Veterinary School of Alfort, Maisons-Alfort, France

Norbert Nowotny

Affiliations Institute of Virology, University of Veterinary Medicine, Vienna, Austria, Department of Microbiology and Immunology, College of Medicine and Health Sciences, Sultan Qaboos University, Muscat, Oman

Jennifer Richardson

Affiliations Virology (UMR1161), French National Institute for Agricultural Research (INRA), Maisons-Alfort, France, Virology (UMR1161), French Agency for Food, Environmental and Occupational Health and Safety (ANSES), Maisons-Alfort, France, Virology (UMR1161), Paris-Est University, National Veterinary School of Alfort, Maisons-Alfort, France

Stéphan Zientara

Affiliations Virology (UMR1161), French National Institute for Agricultural Research (INRA), Maisons-Alfort, France, Virology (UMR1161), French Agency for Food, Environmental and Occupational Health and Safety (ANSES), Maisons-Alfort, France, Virology (UMR1161), Paris-Est University, National Veterinary School of Alfort, Maisons-Alfort, France

Sylvie Lecollinet

Contributed equally to this work with: Sylvie Lecollinet, Muriel Coulpier

Affiliations Virology (UMR1161), French National Institute for Agricultural Research (INRA), Maisons-Alfort, France, Virology (UMR1161), French Agency for Food, Environmental and Occupational Health and Safety (ANSES), Maisons-Alfort, France, Virology (UMR1161), Paris-Est University, National Veterinary School of Alfort, Maisons-Alfort, France

Muriel Coulpier

Contributed equally to this work with: Sylvie Lecollinet, Muriel Coulpier

mcoulpier@vet-alfort.fr

Affiliations Virology (UMR1161), French National Institute for Agricultural Research (INRA), Maisons-Alfort, France, Virology (UMR1161), French Agency for Food, Environmental and Occupational Health and Safety (ANSES), Maisons-Alfort, France, Virology (UMR1161), Paris-Est University, National Veterinary School of Alfort, Maisons-Alfort, France

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ED S. Lecollinet MC. Performed the experiments: ED S. Lowenski TL JLS EL S. Lecollinet. Analyzed the data: ED JLS EL NN JR SZ S. Lecollinet MC. Wrote the manuscript: ED JLS EL NN JR SZ S. Lecollinet MC.