About the Authors

Qing Gao

Contributed equally to this work with: Qing Gao, Zhihua Ju, Yan Zhang

Affiliations Dairy Cattle Research Center, Shandong Academy of Agricultural Science, Jinan, PR China, College of Life Science, Shandong Normal University, Jinan, PR China

Zhihua Ju

Contributed equally to this work with: Qing Gao, Zhihua Ju, Yan Zhang

Affiliation Dairy Cattle Research Center, Shandong Academy of Agricultural Science, Jinan, PR China

Yan Zhang

Contributed equally to this work with: Qing Gao, Zhihua Ju, Yan Zhang

Affiliation Dairy Cattle Research Center, Shandong Academy of Agricultural Science, Jinan, PR China

Jinming Huang

Affiliation Dairy Cattle Research Center, Shandong Academy of Agricultural Science, Jinan, PR China

Xiaojian Zhang

Affiliation Dairy Cattle Research Center, Shandong Academy of Agricultural Science, Jinan, PR China

Chao Qi

Affiliation Dairy Cattle Research Center, Shandong Academy of Agricultural Science, Jinan, PR China

Jianbin Li

Affiliation Dairy Cattle Research Center, Shandong Academy of Agricultural Science, Jinan, PR China

Jifeng Zhong

Affiliation Dairy Cattle Research Center, Shandong Academy of Agricultural Science, Jinan, PR China

Guorong Li

liguorong66@hotmail.com (GL); wangcf1967@163 (CW)

Affiliation College of Life Science, Shandong Normal University, Jinan, PR China

Changfa Wang

liguorong66@hotmail.com (GL); wangcf1967@163 (CW)

Affiliation Dairy Cattle Research Center, Shandong Academy of Agricultural Science, Jinan, PR China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: GL CW. Performed the experiments: QG ZJ YZ JH XZ CQ. Analyzed the data: QG ZJ YZ GL CW. Contributed reagents/materials/analysis tools: JH CQ JL JZ CW. Wrote the paper: QG ZJ YZ GL CW.