About the Authors

Katherine J. Nicholas

Contributed equally to this work with: Katherine J. Nicholas, Emily K. Zern

Affiliation Department of Pathology, Microbiology and Immunology, Department of Medicine, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America

Emily K. Zern

Contributed equally to this work with: Katherine J. Nicholas, Emily K. Zern

Affiliation Division of Infectious Diseases, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America

Louise Barnett

Affiliation Division of Infectious Diseases, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America

Rita M. Smith

Affiliation Division of Infectious Diseases, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America

Shelly L. Lorey

Current address: Department of Cell and Developmental Biology, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America

Affiliation Division of Infectious Diseases, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America

Courtney A. Copeland

Affiliation Department of Molecular Physiology and Biophysics, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America

Shanmugalakshmi Sadagopal

Current address: Yerkes Primate Research Center, Emory University, Atlanta, Georgia, United States of America

Affiliation Division of Infectious Diseases, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America

Spyros A. Kalams

s.kalams@vanderbilt.edu

Affiliations Department of Pathology, Microbiology and Immunology, Department of Medicine, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America, Division of Infectious Diseases, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KJN EKZ SS SAK. Performed the experiments: KJN EKZ RMS SLL CAC SS. Analyzed the data: KJN EKZ LB SAK. Wrote the manuscript: KJN EKZ SAK.