About the Authors

Lisa Y. Pang

lisa.pang@ed.ac.uk

Affiliation Royal (Dick) School of Veterinary Studies and Roslin Institute, The University of Edinburgh, Easter Bush, Midlothian, United Kingdom

Emma L. Gatenby

Affiliation Royal (Dick) School of Veterinary Studies and Roslin Institute, The University of Edinburgh, Easter Bush, Midlothian, United Kingdom

Ayako Kamida

Affiliation Royal (Dick) School of Veterinary Studies and Roslin Institute, The University of Edinburgh, Easter Bush, Midlothian, United Kingdom

Bruce A. Whitelaw

Affiliation Royal (Dick) School of Veterinary Studies and Roslin Institute, The University of Edinburgh, Easter Bush, Midlothian, United Kingdom

Ted R. Hupp

Affiliation Edinburgh Cancer Research UK Centre, The University of Edinburgh, Edinburgh, United Kingdom

David J. Argyle

Affiliation Royal (Dick) School of Veterinary Studies and Roslin Institute, The University of Edinburgh, Easter Bush, Midlothian, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: LP TRH BAW DJA. Performed the experiments: LP ELG AK. Analyzed the data: LP. Contributed reagents/materials/analysis tools: LP. Wrote the paper: LP.