About the Authors

Ryoi Tamura

Contributed equally to this work with: Ryoi Tamura, Hiroshi Nishida

rtamura@med.u-toyama.ac.jp

Affiliation Department of Integrative Neuroscience, Graduate School of Medicine and Pharmaceutical Sciences, University of Toyama, Toyama, Japan

Hiroshi Nishida

Contributed equally to this work with: Ryoi Tamura, Hiroshi Nishida

Affiliation Department of Otorhinolaryngology, Graduate School of Medicine and Pharmaceutical Sciences, University of Toyama, Toyama, Japan

Satoshi Eifuku

Affiliation Department of Systems Neuroscience, School of Medicine, Fukushima Medical University, Fukushima, Japan

Hiroaki Fushiki

Affiliation Faculty of Health Sciences, Mejiro University, Saitama, Japan

Yukio Watanabe

Affiliation Department of Otorhinolaryngology, Graduate School of Medicine and Pharmaceutical Sciences, University of Toyama, Toyama, Japan

Kumiko Uchiyama

Affiliation Department of Integrative Neuroscience, Graduate School of Medicine and Pharmaceutical Sciences, University of Toyama, Toyama, Japan

Competing Interests

The funding from MEXT, JSPS, and Japan Chemical Industry Association do not alter the authors' adherence to all of the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: RT HN. Performed the experiments: RT HN. Analyzed the data: RT HN S.E. Contributed reagents/materials/analysis tools: KU RT. Wrote the paper: RT HN. Designed the software used in analysis: RT.